Puss in Boots discovers that his passion for adventure has taken its toll: He has burned through eight of his nine lives, leaving him with only one life left. Puss sets out on an epic journey to find the mythical Last Wish and restore his nine lives.

Searching for a safer habitat, the prehistoric Crood family discovers an idyllic, walled-in paradise that meets all of its needs. Unfortunately, they must also learn to live with the Bettermans -- a family that's a couple of steps above the Croods on the evolutionary ladder. As tensions between the new neighbors start to rise, a new threat soon propels both clans on an epic adventure that forces them to embrace their differences, draw strength from one another, and survive together.